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The frame is either the time in milliseconds when that value is applied from the start of the effect, or if it’s an “overtime” attribute then it will be a percent of the lifetime of the effect. So for example, if the particle has a lifetime of 3000ms (3 seconds), then a frame of 0.5 will mean that that attributes value will be at 1500ms or 50% into the particles lifetime. Values at any given time are linear interpolated between values, or interpolated with bezier curves if curves are used in the editor.
Yes the animation properties is only for the exporting of the effects and do not affect anything with the attributes.
Hope that helps!